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The Boss 429 engine was derived from the Ford 385 engine. After much consideration, it was decided by Ford that the Mustang would be the car that would house this new engine. NASCAR's Homologation rules required that at least 500 cars be fitted with this motor and sold to the general public.
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Ford was seeking to develop a Hemi engine that could compete with the famed 426 Hemi from Chrysler in NASCAR's Sprint Cup Series (then known as "Grand National Division"). The origin of the Boss 429 comes about as a result of NASCAR. In total there were 1,358 Boss 429's made. The Boss 429 is arguably one of the rarest and most valued Muscle car to date. The Boss 429 was a high performance Ford Mustang variant offered in 19.